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He has not been seen publicly since then, and his current whereabouts and condition are unknown.

To say that Mr. Ahmadinejad was an unusual choice would be a vast understatement. While he had increasingly clashed with the regime’s leaders and had been placed under close watch by the Iranian authorities, he was known during his term as president, from 2005 to 2013, for his calls to “wipe Israel off the map.” He was a strong supporter of Iran’s nuclear program, a fierce critic of the United States, and known for violently cracking down on internal dissent.

How Mr. Ahmadinejad was recruited to take part remains unknown.

The existence of the effort, which has not been previously reported, was part of a multistage plan developed by Israel to topple Iran’s theocratic government. It underscores how Mr. Trump and Prime Minister Benjamin Netanyahu of Israel went into the war not only misjudging how quickly they could achieve their objectives but also gambling to some degree on a risky plan for leadership change in Iran that even some of Mr. Trump’s aides found implausible. Some American officials were skeptical in particular about the viability of putting Mr. Ahmadinejad back into power.

“From the outset, President Trump was clear about his goals for Operation Epic Fury: destroy Iran’s ballistic missiles, dismantle their production facilities, sink their navy, and weaken their proxy,” Anna Kelly, a White House spokeswoman, said in response to a request for comment about the regime change plan and Ahmadinejad. “The United States military met or exceeded all of its objectives, and now, our negotiators are working to make a deal that would end Iran’s nuclear capabilities for good.”

A spokesperson for Mossad, the Israeli foreign intelligence agency, declined to comment.

U.S. officials spoke during the early days of the war about plans developed with Israel to identify a pragmatist who could take over the country. Officials insisted that there was intelligence that some within the Iranian regime would be willing to work with the United States, even if those people couldn’t be described as “moderates.”

Mr. Trump was enjoying the success of the raid by U.S. forces to capture Venezuela’s leader, Nicolas Maduro, and the willingness of his interim replacement to work with the White House — a model that Mr. Trump appeared to think could be replicated elsewhere.

In recent years, Mr. Ahmadinejad has clashed with regime leaders, accusing them of corruption, and rumors have swirled about his loyalties. He was disqualified from numerous presidential elections, his aides were arrested, and Mr. Ahmadinejad’s movements were increasingly restricted to his home in the Narmak section of eastern Tehran.

That American and Israeli officials saw Mr. Ahmadinejad as a potential leader of a new government in Iran is further evidence that the war in February was launched with the hopes of installing more pliable leadership in Tehran. Mr. Trump and members of his cabinet have said that the goals of the war were narrowly focused on destroying Iran’s nuclear, missile, and military capabilities.

There are many unanswered questions about how Israel and the United States planned to put Mr. Ahmadinejad in power, and the circumstances surrounding the airstrike that injured him. American officials said that the strike — carried out by the Israeli Air Force — was meant to kill the guards watching over Mr. Ahmadinejad as part of a plan to release him from house arrest.

On the first day of the war, Israeli strikes killed Ayatollah Ali Khamenei, Iran’s supreme leader. The strike at Ayatollah Khamenei’s compound in central Tehran also blew up a meeting of Iranian officials, killing some officials whom the White House had identified as more willing to negotiate over a change in government than their bosses.

There were also initial reports at the time in the Iranian media that Mr. Ahmadinejad had been killed in the strike on his home.

The strike did not significantly damage Mr. Ahmadinejad’s house at the end of a dead-end street. But the security outpost at the entrance to the street was struck. Satellite imagery shows that building was destroyed.

In the days that followed, official news agencies clarified that he had survived but that his “bodyguards” — in actuality, Islamic Revolutionary Guard Corps members who were both guarding him and holding him under house arrest — were killed.

An article in The Atlantic in March, citing anonymous associates of Mr. Ahmadinejad, said that the former president had been freed from government confinement after the strike at his house, which the article described as “in effect a jailbreak operation.”

After that article, an associate of Mr. Ahmadinejad confirmed to The New York Times that Mr. Ahmadinejad saw the strike as an attempt to free him. The associate said the Americans viewed Mr. Ahmadinejad as someone who could lead Iran, and had the capability to manage “Iran’s political, social, and military situation.”

Mr. Ahmadinejad would have been able to “play a very important role” in Iran in the near future, the associate said, suggesting that the United States saw him as similar to Delcy Rodriguez, who took power in Venezuela after American forces seized Mr. Maduro and has since worked closely with the Trump administration, the person said.

During his presidency, Mr. Ahmadinejad was known both for his hard-line policies and his often outlandish fundamentalist pronouncements, such as his declaration that there was not a single gay person in Iran and his denial of the Holocaust. He spoke at a conference in Tehran called “A World Without Zionism.”

Western satirists lampooned these views, and Mr. Ahmadinejad became something of an unwitting pop culture curiosity, even the subject of Saturday Night Live parodies.

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Former President Mahmoud Ahmadinejad of Iran registering as a candidate in the presidential election in Tehran in 2024. Credit…Arash Khamooshi/Polaris for The New York Times

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